Evolutionary morphology, biomechanics behavior, and systematics of arachnids and other arthropods
Taxonomy, evolution and reproductive biology of leiobunine harvestmen (daddy longlegs) of eastern North America
Hydrodynamics of gill ventilation in mayfly naiads, mechanics of elastic sclerites and hydraulics in arachnid locomotion, molecular systematics of major arthropod groups, comparative anatomy of scorpions and many others